Ward C. Wheeler
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American arachnologist
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Biology
Ward C. Wheeler's Degrees
- PhD Entomology University of California, Davis
- Masters Entomology University of California, Davis
- Bachelors Biology University of California, Davis
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(Suggest an Edit or Addition)Ward C. Wheeler's Published Works
Number of citations in a given year to any of this author's works
Total number of citations to an author for the works they published in a given year. This highlights publication of the most important work(s) by the author
Published Works
- THE AMPHIBIAN TREE OF LIFE (2006) (1912)
- Broad phylogenomic sampling improves resolution of the animal tree of life (2008) (1772)
- SYSTEMATIC REVIEW OF THE FROG FAMILY HYLIDAE, WITH SPECIAL REFERENCE TO HYLINAE: PHYLOGENETIC ANALYSIS AND TAXONOMIC REVISION (2005) (926)
- The Strepsiptera problem: phylogeny of the holometabolous insect orders inferred from 18S and 28S ribosomal DNA sequences and morphology. (1997) (893)
- Assessing the root of bilaterian animals with scalable phylogenomic methods (2009) (710)
- OPTIMIZATION ALIGNMENT: THE END OF MULTIPLE SEQUENCE ALIGNMENT IN PHYLOGENETICS? (1996) (666)
- PHYLOGENETIC SYSTEMATICS OF DART-POISON FROGS AND THEIR RELATIVES (AMPHIBIA: ATHESPHATANURA: DENDROBATIDAE) (2006) (554)
- Arthropod phylogeny based on eight molecular loci and morphology (2001) (527)
- SEQUENCE ALIGNMENT, PARAMETER SENSITIVITY, AND THE PHYLOGENETIC ANALYSIS OF MOLECULAR DATA (1995) (502)
- The Phylogeny of the Extant Hexapod Orders (2001) (490)
- Triploblastic relationships with emphasis on the acoelomates and the position of Gnathostomulida, Cycliophora, Plathelminthes, and Chaetognatha: a combined approach of 18S rDNA sequences and morphology. (2000) (357)
- The Phylogeny of the Extant Chelicerate Orders (1998) (325)
- The spider tree of life: phylogeny of Araneae based on target‐gene analyses from an extensive taxon sampling (2017) (305)
- On bivalve phylogeny: a high‐level analysis of the Bivalvia (Mollusca) based on combined morphology and DNA sequence data (2005) (299)
- ARTHROPOD PHYLOGENY: A COMBINED APPROACH (1993) (290)
- Multiple sequence alignment in phylogenetic analysis. (2000) (263)
- Paired sequence difference in ribosomal RNAs: evolutionary and phylogenetic implications. (1988) (252)
- NUCLEIC ACID SEQUENCE PHYLOGENY AND RANDOM OUTGROUPS (1990) (246)
- On gaps. (1999) (244)
- Phylogenomic interrogation of arachnida reveals systemic conflicts in phylogenetic signal. (2014) (234)
- The position of arthropods in the animal kingdom: Ecdysozoa, islands, trees, and the "Parsimony ratchet". (1999) (212)
- Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing (2005) (211)
- Venom evolution widespread in fishes: a phylogenetic road map for the bioprospecting of piscine venoms. (2006) (209)
- POY version 4: phylogenetic analysis using dynamic homologies (2010) (207)
- Phylogeny and Systematic Position of Opiliones: A Combined Analysis of Chelicerate Relationships Using Morphological and Molecular Data 1 (2002) (196)
- Taxonomic Impediment or Impediment to Taxonomy? A Commentary on Systematics and the Cybertaxonomic-Automation Paradigm (2007) (194)
- Phylogenetic relationships among superfamilies of Hymenoptera (2012) (184)
- Implied alignment: a synapomorphy‐based multiple‐sequence alignment method and its use in cladogram search (2003) (183)
- Evolution of the hymenopteran megaradiation. (2011) (180)
- Cladistic relationships among higher groups of Heteroptera: congruence between morphological and molecular data sets (1993) (177)
- Phylogenetic relationships within the Cimicomorpha (Hemiptera: Heteroptera): a total‐evidence analysis (2009) (170)
- Elision: a method for accommodating multiple molecular sequence alignments with alignment-ambiguous sites. (1995) (159)
- MALIGN: A Multiple Sequence Alignment Program (1994) (155)
- Systematics of the Damon variegatus group of African whip spiders (Chelicerata: Amblypygi): Evidence from behaviour, morphology and DNA (2005) (135)
- Polyphyly of the mail-cheeked fishes (Teleostei: Scorpaeniformes): evidence from mitochondrial and nuclear sequence data. (2004) (133)
- Techniques in Molecular Systematics and Evolution (2002) (131)
- Simultaneous analysis of the basal lineages of Hymenoptera (Insecta) using sensitivity analysis (2002) (128)
- Dynamic homology and phylogenetic systematics: a unified approach using POY (2006) (126)
- Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera) (2008) (125)
- The phylogenetic relationships of the charismatic poster frogs, Phyllomedusinae (Anura, Hylidae) (2009) (124)
- Homology and the Optimization of DNA Sequence Data (2001) (116)
- Fixed Character States and the Optimization of Molecular Sequence Data (1999) (115)
- Phylogeny of the sea spiders (Arthropoda, Pycnogonida) based on direct optimization of six loci and morphology (2007) (114)
- Phylogeny of Henicopidae (Chilopoda: Lithobiomorpha): a combined analysis of morphology and five molecular loci (2002) (112)
- Phylogeny of Branchiopoda (Crustacea) based on a combined analysis of morphological data and six molecular loci (2007) (104)
- The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow‐mouthed frogs (Anura, Microhylidae) (2016) (104)
- The amphibian tree of life. Bulletin of the AMNH ; no. 297 (2006) (103)
- Higher level relationships of the arctoid Carnivora based on sequence data and "total evidence". (1994) (100)
- Mitochondrial DNA Size Variation Within Individual Crickets (1985) (100)
- Systematics and biogeography of the family Scorpionidae (Chelicerata : Scorpiones), with a discussion on phylogenetic methods (2003) (93)
- COMBINATORIAL WEIGHTS IN PHYLOGENETIC ANALYSIS: A STATISTICAL PARSIMONY PROCEDURE (1990) (92)
- Molecular Systematics and Evolution: Theory and Practice (2002) (92)
- Systematics of spiny‐backed treefrogs (Hylidae: Osteocephalus): an Amazonian puzzle (2013) (90)
- INDIVIDUAL ORGANISMS AS TERMINAL ENTITIES: LAYING THE SPECIES PROBLEM TO REST (1992) (88)
- Is Ellipura monophyletic? A combined analysis of basal hexapod relationships with emphasis on the origin of insects (2004) (85)
- Insect homeotic transformation (1994) (82)
- A molecular perspective on the phylogeny of the Hyla pulchella species group (Anura, Hylidae). (2004) (81)
- POY version 5: phylogenetic analysis using dynamic homologies under multiple optimality criteria (2015) (76)
- Genomic analysis and geographic visualization of the spread of avian influenza (H5N1). (2007) (72)
- Linking of digital images to phylogenetic data matrices using a morphological ontology. (2007) (71)
- A phylogenetic analysis of Pleurodema (Anura: Leptodactylidae: Leiuperinae) based on mitochondrial and nuclear gene sequences, with comments on the evolution of anuran foam nests (2012) (71)
- Into the deep: a phylogenetic approach to the bivalve subclass Protobranchia. (2013) (69)
- Phylogenetic relationships of a Patagonian frog radiation, the Alsodes + Eupsophus clade (Anura: Alsodidae), with comments on the supposed paraphyly of Eupsophus (2013) (69)
- Comparative and phylogenetic analysis of developmental sequences (2004) (67)
- Revisiting habitat and lifestyle transitions in Heteroptera (Insecta: Hemiptera): insights from a combined morphological and molecular phylogeny (2019) (66)
- Systematics: A Course of Lectures (2012) (65)
- Some Unusual Small-Subunit Ribosomal RNA Sequences of Metazoans (2001) (64)
- Iterative pass optimization of sequence data (2003) (63)
- The evolutionary transition from subsocial to eusocial behaviour in Dictyoptera: phylogenetic evidence for modification of the "shift-in-dependent-care" hypothesis with a new subsocial cockroach. (2007) (62)
- Sampling, groundplans, total evidence and the systematics of arthropods (1998) (60)
- Iterative pass optimization of sequence data. (2003) (54)
- Hox gene duplications correlate with posterior heteronomy in scorpions (2014) (52)
- Dynamic homology and the likelihood criterion (2006) (50)
- Measuring Topological Congruence by Extending Character Techniques (1999) (48)
- Sources of ambiguity in nucleic acid sequence alignment. (1994) (44)
- Phylogeny, Taxonomic Revision, and Character Evolution of the Genera Chiasmocleis and Syncope (Anura, Microhylidae) in Amazonia, with Descriptions of Three New Species (2014) (43)
- Maximum Parsimony on Phylogenetic networks (2012) (42)
- Revenant clades in historical biogeography: the geology of New Zealand predisposes endemic clades to root age shifts (2013) (39)
- Dissociation between hand motion and population vectors from neural activity in motor cortex (36)
- Systematics of Polistes (Hymenoptera: Vespidae), with a phylogenetic consideration of Hamilton's haplodiploidy hypothesis (2006) (36)
- Phylogenetic network analysis as a parsimony optimization problem (2015) (36)
- 13 – Homology and DNA Sequence Data (2001) (36)
- Regular ArticleArthropod Phylogeny: a Combined Approach (1993) (36)
- Optimization Alignment:Down,Up,Error,and Improvements (2002) (36)
- Homology and the optimization of DNA sequence data. (2001) (34)
- DNA multiple sequence alignments. (2002) (34)
- Is The Amphibian Tree of Life really fatally flawed? (2008) (32)
- On the Monophyly and Relationships of Several Genera of Hylini (Anura: Hylidae: Hylinae), with Comments on Recent Taxonomic Changes in Hylids (2018) (32)
- First global molecular phylogeny and biogeographical analysis of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid‐Cretaceous diversification (2017) (31)
- MISSING ENTRY REPLACEMENT DATA ANALYSIS: A REPLACEMENT APPROACH TO DEALING WITH MISSING DATA IN PALEONTOLOGICAL AND TOTAL EVIDENCE DATA SETS (2003) (31)
- Efficiency of parallel direct optimization. (2001) (30)
- Alignment characters, dynamic programming and heuristic solutions (1998) (30)
- The tree alignment problem (2012) (29)
- Topology-Bayes versus Clade-Bayes in phylogenetic analysis. (2008) (28)
- Poor taxon sampling, poor character sampling, and non‐repeatable analyses of a contrived dataset do not provide a more credible estimate of insect phylogeny: a reply to Kjer (2005) (28)
- A conserved genetic mechanism specifies deutocerebral appendage identity in insects and arachnids (2015) (27)
- Phylogenomic Resolution of Sea Spider Diversification through Integration of Multiple Data Classes (2020) (26)
- Molecular phylogeny of Indo‐Pacific carpenter ants (Hymenoptera: Formicidae, Camponotus) reveals waves of dispersal and colonization from diverse source areas (2015) (26)
- Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. (2014) (24)
- The Supramap project: linking pathogen genomes with geography to fight emergent infectious diseases (2011) (24)
- Alignment, dynamic homology, and optimization (2006) (24)
- Taxon cycle predictions supported by model‐based inference in Indo‐Pacific trap‐jaw ants (Hymenoptera: Formicidae: Odontomachus) (2018) (23)
- Evidence of duplicated Hox genes in the most recent common ancestor of extant scorpions (2015) (23)
- Evolution in the Genus Rhinella: A Total Evidence Phylogenetic Analysis of Neotropical True Toads (Anura: Bufonidae) (2021) (22)
- Cross-bracing uncalibrated nodes in molecular dating improves congruence of fossil and molecular age estimates (2014) (22)
- 'Pluralism' and the aims of phylogenetic research. (2002) (22)
- Parsimony overcomes statistical inconsistency with the addition of more data from the same gene (2005) (22)
- Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia, Athesphatanura, Dendrobatidae). Bulletin of the AMNH ; no. 299 (2006) (21)
- Systematic review of the frog family Hylidae, with special reference to Hylinae : phylogenetic analysis and taxonomic revision. Bulletin of the AMNH ; no. 294 (2005) (21)
- The case for sensitivity: a response to Grant and Kluge (2007) (21)
- Search‐based optimization (2003) (21)
- Local search for the generalized tree alignment problem (2013) (20)
- A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) and parametric shape analysis reveal the disutility of subfamilial nomenclature in armored harvestman systematics. (2017) (20)
- POY version 3.0.11 (Wheeler, Gladstein and De Laet, May 6 2003). Command line documentation (2003) (20)
- Historical linguistics as a sequence optimization problem: the evolution and biogeography of Uto‐Aztecan languages (2015) (20)
- Phylogeography of the harvestman genus Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox (2016) (19)
- Subdivision of arthropod cap-n-collar expression domains is restricted to Mandibulata (2014) (19)
- Partition‐free congruence analysis: implications for sensitivity analysis (2006) (19)
- Phylogenetic analysis of the endemic New Caledonian cockroach Lauraesilpha. Testing competing hypotheses of diversification (2008) (17)
- Hennig's semaphoront concept and the use of ontogenetic stages in phylogenetic reconstruction (2017) (17)
- Theory and practice of parallel direct optimization. (2002) (15)
- Molecular evolution and phylogenetic utility of the polyubiquitin locus in mammals and higher vertebrates. (1996) (15)
- A phylogeny of sand flies (Diptera: Psychodidae: Phlebotominae), using recent Ethiopian collections and a broad selection of publicly available DNA sequence data (2015) (14)
- Forked Tongues Revisited: Molecular Apomorphies Support Morphological Hypotheses of Squamate Evolution (2015) (13)
- Expression and function of spineless orthologs correlate with distal deutocerebral appendage morphology across Arthropoda. (2017) (13)
- Trivial minimization of extra‐steps under dynamic homology (2012) (12)
- Elongation factor-1α, a putative single-copy nuclear gene, has divergent sets of paralogs in an arachnid. (2013) (12)
- Comprehensive Species Sampling and Sophisticated Algorithmic Approaches Refute the Monophyly of Arachnida (2021) (11)
- Comparison of heuristic approaches to the generalized tree alignment problem (2016) (11)
- Myrmecicultoridae, a New Family of Myrmecophilic Spiders from the Chihuahuan Desert (Araneae: Entelegynae) (2019) (11)
- Phylogenetic Hypotheses and the Utility of Multiple Sequence Alignment (2007) (11)
- Phylogeny of Map Tree Frogs, Boana semilineata Species Group, with a New Amazonian Species (Anura: Hylidae) (2018) (11)
- Indel information eliminates trivial sequence alignment in maximum likelihood phylogenetic analysis (2012) (10)
- Phylogeny Reconstruction via Direct Optimization of DNA and other data (2002) (9)
- Distinctions between optimal and expected support (2010) (9)
- Preliminary phylogenetic analysis of the Andean clade and the placement of new Colombian blueberries (Ericaceae, Vaccinieae) (2015) (9)
- Pairwise alignment with rearrangements. (2006) (9)
- Nomenclatural stability does not justify recognition of paraphyletic taxa: A response to Scherz et al. (2016). (2017) (9)
- Search-based optimization (2003) (8)
- A NOVEL METHOD FOR ECONOMICAL DIAGNOSIS OF CLADOGRAMS UNDER SANKOFF OPTIMIZATION (1994) (8)
- Application note: on extension gap in POY version 3 (2008) (8)
- Mitochondrial Intergenic Spacer in Fairy Basslets (Serranidae: Anthiinae) and the Simultaneous Analysis of Nucleotide and Rearrangement Data (2009) (7)
- Phylogenetic relationships of the Boana pulchella Group (Anura: Hylidae). (2020) (7)
- Revising the Bantu tree (2018) (7)
- Direct Optimization, Sensitivity Analysis, and the Evolution of the Hymenopteran Superfamilies (2013) (7)
- Towards improving searches for optimal phylogenies. (2015) (6)
- Chromosomal character optimization. (2007) (6)
- Analysis and visualization of H7 influenza using genomic, evolutionary and geographic information in a modular web service (2012) (5)
- Venation patterns of neotropical blueberries (Vaccinieae: Ericaceae) and their phylogenetic utility (2013) (5)
- Exactly Computing the Parsimony Scores on Phylogenetic Networks Using Dynamic Programming (2014) (5)
- Phylogeny, taxonomic revision, and character evolution of the genera Chiasmocleis and Syncope (Anura, Microhylidae) in Amazonia, with descriptions of three new species. (Bulletin of the American Museum of Natural History, no. 386) (2014) (5)
- 6. Phylogenetic Hypotheses and the Utility of Multiple Sequence Alignment (2019) (4)
- Phyletic groups on networks (2014) (4)
- Statistical Modeling of Distribution Patterns: a Markov Random Field Implementation and its Application on Areas of Endemism. (2019) (4)
- An E!cient Heuristic for the Tree Alignment Problem (2008) (4)
- Towards Phylogenomic Reconstruction (2007) (3)
- Molecular data in systematics: a promise fulfi lled, a future beckoning (2016) (3)
- Efficiency of Parallel Direct Optimization (2001) (2)
- Maximum a posteriori probability assignment (MAP‐A): an optimality criterion for phylogenetic trees via weighting and dynamic programming (2014) (2)
- scorpions Hox gene duplications correlate with posterior heteronomy in Supplementary data tml (2014) (1)
- Trait-dependent biogeography: model-based inference of dispersal and distribution patterns of Indo-Pacific trap-jaw ants (Hymenoptera: Formicidae: Odontomachus) (2019) (1)
- Subdivision of arthropod cap-n-collar expression domains is restricted to Mandibulata (2014) (1)
- Cross-bracing uncalibrated nodes in molecular dating improves congruence of fossil and molecular age estimates (2014) (1)
- Comparing and displaying phylogenetic trees using edge union networks (2019) (1)
- Phylogenetic supergraphs (2021) (1)
- There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). (2017) (1)
- Local search for the generalized tree alignment problem (2013) (1)
- REVIEW Multiple Sequence Alignment in Phylogenetic Analysis (2000) (1)
- Algorithms and hypothesis selection in dynamic homology phylogenetic analysis (2010) (0)
- All hail King Ludd (2021) (0)
- FASTC: a file format for multi‐character sequence data (2019) (0)
- Some Clarifications on Phylogenetic Sequence Analysis , Dynamic Homology , Structural Alignment , and the use of POY : A Response to Kjer et al . 2007 (2007) (0)
- Efficient implied alignment (2020) (0)
- Supplemental Material for 'Evolution in the genus Rhinella : a total evidence phylogenetic analysis of neotropical true toads (Anura: Bufonidae). (Bulletin of the American Museum of Natural History, no. 447)' (2021) (0)
- Efficient implied alignment (2020) (0)
- Consensus, Congruence, and Supertrees (2012) (0)
- Optimality Criteria–Posterior Probability (2012) (0)
- Parallel computer processing in systematics (2004) (0)
- Maximum Parsimony on Phylogenetic networks (2012) (0)
- TR-2008015: An Efficient Heuristic for the Tree Alignment Problem (2008) (0)
- Statistical and Mathematical Basics (2012) (0)
- A tale of two bellies: systematics of the oval frogs (Anura: Microhylidae: Elachistocleis Parker, 1927) (2022) (0)
- Continuation of Novel Analytical and Empircal Approaches to the Origin and Prediction of Pathogenicity (2014) (0)
- Reconstructing the Protostome Tree of Life-the past, the present and the future (2010) (0)
- scorpionsgene duplications correlate with posterior heteronomy in (2014) (0)
- Phylogenetic network analysis as a parsimony optimization problem (2015) (0)
- Optimality Criteria–Likelihood (2012) (0)
- COMPUTATIONAL ASPECTS OF THE PHYLOGENETIC ANALYSIS OF COMPARATIVE SEQUENCE DATA (2016) (0)
- Hypothesis Testing and the Philosophy of Science (2012) (0)
- Optimality Criteria–Distance (2012) (0)
- Evidence for reduced BRCA2 functional activity in Homo sapiens after divergence from the chimpanzee-human last common ancestor. (2022) (0)
- The tree alignment problem (2012) (0)
- Selection for Decreased BRCA2 Functional Activity in Homo sapiens After Divergence from the Chimpanzee-Human Last Common Ancestor (2020) (0)
- Distance Wagner tree refinement as a heuristic approach to character‐based initial tree construction (2021) (0)
- Deuterostome Evolution: Large Data Set Analysis (2004) (0)
- Appendix A: Mathematical Notation (2012) (0)
- Species Concepts, Definitions, and Issues (2012) (0)
- Clocks and Rates (2012) (0)
- Phylogeography of the harvestman genus Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox (2015) (0)
- Optimality Criteria–Parsimony (2012) (0)
- Comparison of Optimality Criteria (2012) (0)
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